Fresh out of Glasgow Veterinary College, James Herriot (Nicholas Ralph) follows his dream to become a vet in the magnificent Yorkshire Dales. He soon discovers that treating the animals is as much about treating their owners, and the Dales farmers are a tough crowd to please. Settling in at Skeldale House, James gets to know his newly formed dysfunctional family; his chaotic and erratic boss Siegfried Farnon (Samuel West), his wayward brother Tristan (Callum Woodhouse) and the shrewd housekeeper Mrs. Hall (Anna Madeley) who is endlessly steering the ship. As local farmer's daughter Helen Alderson (Rachel Shenton) wins James's heart he soon finds himself falling in love with more than just the Dales. As Helen and James navigate their feelings for one another, a trip back to Glasgow in Series Two presents a dilemma which leaves James having to decide whether to follow his head or his heart. Series Three brings with it big changes as James enters a new stage of his life, both with Helen and with the practice. With the added potential of looming war, our Skeldale family are forced to consider their purpose in Darrowby and beyond.